What do the different readings of 髑髏 mean?

I recently came across the word 髑髏, but when I looked it up it seemed to have maybe four different readings. Would anyone know what the difference is between them? I’ve been trying to search to no avail.

The standard word is どくろ. I doubt that kanji is going to appear very often.

Everything else looks like irregular / nonstandard readings to me

https://en.wiktionary.org/wiki/髑髏

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